Springboks coach Heyneke Meyer has made five changes to his starting XV. Amongst the backs, there are three changes with Pat Lambie and Cobus Reinach replacing Handré Pollard and Francois Hougaard at fly-half and scrum-half respectively while JP Pietersen comes in for Cornal Hendricks on the right wing. Up front, Schalk Burger takes over from Teboho Mohoje at blindside flank and Adriaan Strauss is preferred to Bismarck du Plessis at hooker.
Bath winger Anthony Watson will make his first start for England in place of the injured Semesa Rokoduguni against South Africa. Elsewhere in the starting XV, Dylan Hartley and Courtney Lawes (both Northampton Saints) successfully completed the graduated return to play protocol.
Hello and welcome to our live coverage of England versus South Africa. Twickenham is the setting for this November international and with both nations entering on the back of losses to New Zealand and Ireland respectively, a great deal rests on the result. Stuart Lancaster and Heyneke Meyer will be desperate not to suffer successive end-of-year defeats in what should be a tight 80 minutes. Team news is coming up.
Here we go! England kick off. The Boks gather without problem and Lambie clears to near halfway
Strong start from England as they gather the line-out and start the first attack but Hartley is caught in possession on the deck. Lambie sets up the line-out.
Opportunity lost for SA. The line-out is won and it goes to the backline. Serfontein gets a dodgy pass but holds on. The ball goes loose and the Boks go offside so Farrell kicks for the touchline this time.
Strong run from Eastmond into the Bok 22 but he losses it in contact with Serfontein. We'll have the first scrum. Solid scrum from the Boks and Reinach clears under pressure.
Billy Vunipola drops it in contact on the charge after the line-out. Another scrum then some kick tennis. Matfield calls for one...but drops it! It's obviously slippery out there.
PENALTY, SOUTH AFRICA, LAMBIE. 0-3. But now Farrell gifts South Africa a chance of points, as instead of clearing from his 22, he ran it. Wilson was the offender at the ruck.
England look for a response but Vunipola knocks on in the tackle again. Le Roux puts a kick deep in the 22 and gets it back. He runs and finds Burger but he is tackled into touch.
TRY, SOUTH AFRICA, SERFONTEIN. Terrible mistake from Danny Care, who takes an age to make his pass. Serfontein intercepts near halfway and bolts clear to score under the posts.
CONVERSION, LAMBIE. 0-10. England face a real test of character if they hope to recover from this now.
Brown with a great take in the air and he races into the 22 but Reinach hunts him down. England regroup but Coetzee gets in over the ball and wins a penalty at the breakdown. Lambie clears to relieve the pressure.
Very strong scrum from England as Marler wins the penalty. Farrell sets up the line-out deep in SA territory.
Barritt takes it up and they win quick ball. Vunipola takes it up to the 22 but it is won on the ground by Reinach. Lambie's kick is charged down but Vermeulen secures it. It is eventually scrambled away by De Villiers to bring some relief.
Burger getting some attention from the medics after taking a bash to the head.
Attwood breaks after the Boks bobble it forward off a boot at a ruck. It comes wide to Eastmond then May, who is stopped just short. They go off their feet but Serfontein had knocked on in the tackle so England have an attacking scrum in the 22.
Another string attack from England but the Boks stand strong. Vunipola takes it off the back but Reinach wraps him up and the Boks get it back. Reinach''s clearance is charged and England have it back. Vermeulen is penalised on the deck. Farrell will line it up.
PENALTY, ENGLAND, FARRELL. 3-10. Easy kick for Farrell.
Penalty from long range but the Boks choose the line-out. Lambie tries a cross-kick but it''s too deep and Brown marks it easily. He finds touch short of halfway so the Boks stay on the attack. Strong maul from South Africa as they rumble forward. Lawes pulls it down - penalty! Lambie lines it up.
PENALTY, SOUTH AFRICA, LAMBIE. 3-13. Solid kick from Lambie, who restores the 10-point lead.
PENALTY, ENGLAND, FARRELL. 6-13. Vermeulen penalised on the floor again. Off his feet. Farrell sends it straight down the middle.
Wood takes Etzebeth out in the air at a line-out. That's a penalty. Lambie will have a crack, 15m in from the right hand touchline.
MISSED penalty from Lambie. England restart. Vermeulen takes it over halfway. Le Roux kicks and Brown is there. England run from deep and go over halfway before Brown kicks. Le Roux finds touch near halfway.
ENGLAND 6-13 SOUTH AFRICA
A nothing kick from Farrell is easily collected by Habana, who takes the mark. That's half time. The Boks lead by 10, but neither side have been particularly good. Less errors from SA. We'll be back in ten.
TRY, SOUTH AFRICA, REINACH. 6-18. Lambie chips over the top and Le Roux collects it on the volley, he races 20 before offloading to Reinach, who finishes! Great try.
Here we go for the second half. Lambie goes deep. Vunipola collects. Farrell clears.
CONVERSION, LAMBIE. 6-20. Easy conversion.
Brown runs it back well and takes it up to the 22 where the Boks go offside. England kick into the corner from the penalty. Great chance to strike back.
YELLOW CARD, SOUTH AFRICA, MATFIELD. Pulling down the maul.
TRY, ENGLAND, WILSON. 11-20. An unstoppable maul from England as the home pack rumbles over.
CONVERSION, FARRELL. 13-20.
Terrible mistake from the officials, who award a line-out to England even though Habana had a foot in touch when he caught the ball.
TRY, ENGLAND, MORGAN. 18-20. Another huge maul from England rumbles forward. They move into the 22 with advantage. Morgan breaks off and muscles his way over!
CONVERSION, FARRELL. 20-20. We're all square, huge comeback from England.
England with their tails up as Brown breaks out again but Etzebeth wins a vital penalty at the breakdown so the Boks can clear it. Lambie finds touch from the penalty. Matfield still in the bin.
Etzebeth claims the line-out and they drive the maul. England go off their feet again. Penalty. Lambie goes for the corner again.
TRY, SOUTH AFRICA, BURGER. 20-25. Another great maul results in Burger going over in the corner. Not bad work without Matfield!
Lambie MISSES the conversion. 20-25 it remains. Matfield returns to the fray.
Farrell's restart goes straight out! The Boks drive another maul from halfway with a full pack. Lambie lofts one into the sky, Watson knocks on as he tries to collect. Burger is on hand to pounce but doesn't gather cleanly. Scrum to SA on the England 22.
De Villiers takes it up in midfield and it is quick ball in the 22. The forwards take over and they bash away.
Lambie tries a cross-field kick for Habana to chase. Watson is under it and is taken out in the air. Penalty, but it's off the side of Farrell's boot.
YELLOW CARD, ENGLAND, HARTLEY. Stupid, stupid, stupid. The England hooker stamps on Vermeulen, who is lying on the ground. Walsh has a look on the screen and brings out the card.
Lambie fluffs the kick for the corner, sending it over the deadball line! Horrible mistake. Then the Boks concede a penalty to a 7-man England scrum.
Trouble here for Jannie du Plessis, who has a knee injury. He's off. Coenie Oosthuizen comes on. Ben Youngs and George Ford replace Owen Farrell and Danny Care. Bakkies Botha replaces Eben Etzebeth.
Another kick goes off the side of Farrell's boot. He's having a bad game. England are ready to change their halfbacks.
PENALTY, SOUTH AFRICA, LAMBIE. 20-28. Straight down the middle from Lambie from long range after Attwood is penalised for interfering with Botha in the air at the line-out.
PENALTY, ENGALND, FORD. 23-28. Botha penalised for holding on on the ground.
Brown does well to pluck one out of the air under pressure from Habana. Watson makes good ground, so does Robshaw. England have their tails up. It breaks down but we'll come back for a penalty against Coetzee for a high tackle. Ford kicks into the corner.
England maul it but the loose ball is picked up by Habana and the Boks can counter. Loose pass from Le Roux but they secure it and Reinach clears. Le Roux takes Brown in the air so England kick to the corner again. Match in the balance here.
Line-out steal! Vermeulen at the tail...how crucial could that be? Lambie clears. Ford kicks it out on the full!
Another Bok line-out, another maul. Lawes penalised - that seems harsh. Lambie kicks into the corner.
Ferocious contest as the Bok big men hammer away, just short of the England line. A try here will secure the match...
DROP-GOAL, SOUTH AFRICA, LAMBIE. 23-31. Clinical!
TRY, ENGLAND, BARRITT. 28-31. England leave the best for last, but is it too late? Great try finished by Barritt in the corner off May's pass.
Ford MISSES the conversion but there IS time for the restart.
ENGLAND 28-31 SOUTH AFRICA
Extra time now as England look to come from their own 22. It is lost forward and the Boks keep playing. Has Willie le Roux scored off Reinach's chip ahead? The TMO says no! It doesn't matter, the Springboks are the winners anyway. Thanks for joining us, cheers!